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For full text PDFs of no OA publications, please, send an email by contacts menu or visit the ResearchGate profile of Ivan Nunes (PI):

Manuscripts Submitted or Accepted for Publication

(1) TOLEDO, L. F. ; BOTELHO, L. M. ; CARRASCO-MEDINA, A. S. ; GRAY, J. A. ; ERNETTI, J. R. ; GAMA, J. M. ; LYRA, M. L. ; BLACKBURN, D. C. ; NUNES, I. ; MUSCAT, E. . Among the world's smallest vertebrates: a new miniaturized Flea-Toad (Brachycephalidae) from the Atlantic rainforest. PeerJ, 2024. (Accepted)

(2) Zootaxa, 2024. (subm.)

(3) Biota Neotropica, 2024 (subm.)

(4) Biological Invasions, 2023 (subm.)

(5) Herpetology Notes, 2023. (subm.)

(6) Scientific Reports, 2023. (subm.)

Peer-reviewed Articles

44) VELASCO, I.G. & NUNES, I. Testing the efficiency of passive acoustic monitoring and active monitoring in anuran community in lotic environments in Itinguçu State Park, Peruíbe-SP. Anais da Academia Brasileira de Ciências, v.96, p. e20231057, 2024.

43) RIVERA-CORREA, M.; RON, S. R. ; NUNES, I.; ARAUJO-VIEIRA, K.; PINHEIRO, P. D. P.; GRANT, T. Forty years later: a new Andean stream treefrog of the genus Hyloscirtus (Anura: Hylidae) from Ecuador, with comments on arm hypertrophy in the H. larinopygion group. Zootaxa, v.5474, p. 101-126, 2024.


42) ARAUJO-VIEIRA, K.; LOURENCO, A. C. C.; LACERDA, J. V.; LYRA, M. L.; BLOTTO, B. L.; Ron, S. R.; BALDO, D.; PEREYRA, M. O.; SUAREZ-MAYORGA, A.; BAÊTA, D.; FERREIRA, R. B.; BARRIO-AMOROS, C. L.; BORTEIRO, C.; BRANDAO, R. A.; BRASILEIRO, C. A.; DONNELLY, M. A.; DUBEUX, M. J. M.; KOHLER, J.; KOLENC, F.; LEITE, F. S. F.; MACIEL, N. M.; NUNES, I.; ORRICO, V. G. D.; PELOSO, P. L. V.; PEZZUTI, T. L. , et al. ; Treefrog diversity in the Neotropics: Phylogenetic relationships of Scinaxini (Anura: Hylidae: Hylinae). South American Journal of Herpetology, v. 27, p. 1-143, 2023.


41) MOROTI, M. T.; PEDROZO, M.; SEVERGNINI, M. R.; AUGUSTO-ALVES, G.; DENA, S.; MARTINS, I.; NUNES, I.; MUSCAT, E. A new species of Odontophrynus (Anura, Odontophrynidae) from the southern portion of the Mantiqueira mountains. European Journal of Taxonomy, v. 847, p. 160-193, 2022.


40) COSTA, F. R. ; MOURA, P. H. A. G. ; ALVES JUNIOR, S. R.; ROSA, P. S.; NUNES, I. Acoustic analysis of vocalization and the behavioral response associated to sound production of nine-banded armadillos (Dasypus novemcinctus) in an agonistic context. Papéis Avulsos de Zoologia, v. 62, p. e202262018, 2022.

39) NUNES, I.; GUIMARAES, C. S.; MOURA, P. H. A. G.; PEDROZO, M.; MOROTI, M. T.; CASTRO, L. M.; STUGINSKI, D.R.;MUSCAT, E. Hidden by the name: a new fluorescent pumpkin toadlet from the Brachycephalus ephippium group (Anura: Brachycephalidae). PLoS One, v. 16, p. e0244812, 2021.

38) MOURA, P. H. A. G.; ELIAS-COSTA, A. J.; NUNES, I.; FAIVOVICH, J. Diversity and Evolution of the Extraordinary Vocal Sacs of Casque-Headed Treefrogs (Anura: Hylidae). Biological Journal of the Linnean Society. v. 134, p. 423-442, 2021.

The phylogeny of the Casque‐headed Treefrogs (Hylidae: Hylinae: Lophyohylini). Cladistics, v. 37, p. 36-72, 2021.

36) BECCARI, V.; PINHEIRO, F. L.; NUNES, I.; ANELLI, L. E.; MATEUS, O.; COSTA, F. R. Osteology of an exceptionally well-preserved tapejarid skeleton from Brazil: revealing the anatomy of a curious pterodactyloid clade. PLoS One, v. 16, p. e0254789, 2021.

35) MOURA, P. H. A. G.; COSTA, F. R.; ANELLI, L. E.; NUNES, I. A new genus of fossil frog (Anura) from lower Cretaceous deposits in South America. Anais da Academia Brasileira de Ciênicas, v. 93 (Suppl.2), p. e20191560, 2021.

34) SICHIERI, G. R. F.; CRUZ, C. A. G.; PIMENTA, B. V. S.; NUNES, I. Advertisement call description of two Proceratophrys species (Anura: Odontophrynidae). Zootaxa, v. 4975, p. 397-400, 2021.

33) FERNANDES, D. L.; NUNES, I.; COSTA, F. R. A taxonomic approach on diagnostic characters used to define new pterosaur taxa and an estimation of pterosaur diversity. Anais da Academia Brasileira de Ciênicas, v. 93, p. e20201568, 2021.

32) SICHIERI, G. R. F.; LOPES, B. S.; MOURA, P. H. A. G.; FACIOLE, F. J.; COSTA, F. R.NUNES, I. Predation on leptodactylid and hylid frogs in the Brazilian Pantanal by the invasive Dilocarcinus pagei Stimpson, 1861 (Decapoda, Trichodactylidae). Herpetology Notes, v. 14, p. 1133-1136, 2021.

31) COSTA, F. R. ; MOURA, P. H. A. G. ; NUNES, I. . On the courtship, breeding behaviour and vocalization of Rhinella ornata (Spix, 1824) (Anura, Bufonidae): a well-marked escalated behaviour in a lek-like system. Acta Ethologica, v. 2020, p. 1-9, 2020.

30) SICHIERI, G. R. F.; MOROTI, M. T.; COSTA, F. R.; MUSCAT, E.; NUNES, I. Redescription of the tadpole of rare Giant Torrent Frog Megaelosia boticariana Giaretta & Aguiar, 1998 (Anura, Hylodidae) with comments on natural history. Zootaxa, v. 4896, p. 140-144, 2020.

29) ANTONIO, K. L. R.; SICHIERI, G. R. F.; MOURA, P. H. A. G.; SAMELO, R. R.; COSTA, F. R.; NUNES, I. Caught in the alien web: predation of exotic Anolis porcatus Gray, 1840 (Squamata: Dactyloidae) by the exotic Cyrtophora citricola (Forskål, 1775) (Araneae: Araneidae) in Brazil. Herpetology Notes, v.13, p.1099-1101, 2020.

28) MUSCAT, E.; STUGINSKI, D.; NUNES, I.; MARTINS, I.; AUGUSTO-ALVES, G.; TOLEDO, L. F.; MOROTI, M. T. Update on the geographic distribution of three poorly known frog species in the Mantiqueira mountain range. Herpetology Notes. v. 13, p. 573-577, 2020.

27) MOURA, P. H. A. G. ; ALVES JUNIOR, S. R. ; SOUSA, D. G. S. E. ; CORREA, C. N. ; NUNES, I. . Redescription of the tadpole of Thoropa taophora (Miranda-Ribeiro, 1923) (Anura: Cycloramphidae). Zootaxa, v. 4656, p. 397-400, 2019.

26) ORRICO, V. G. D. ; NUNES, I. ; MATTEDI, C. ; FOUQUET, A. ; LEMOS, A.W. ; RIVERA-CORREA, M. ; LYRA, M. L. ; LOEBMANN, D. ; PIMENTA, B. V. S. ; CARAMASCHI, U. ; RODRIGUES, M. T. ; HADDAD, C. F. B. . Integrative taxonomy supports the existence of two distinct species within Hypsiboas crepitans (Anura: Hylidae). Salamandra, v. 53, p. 99-113, 2017.

25) NUNES, I.; LOEBMANN, D. ; CRUZ, C. A. G. ; HADDAD, C. F. B. . Advertisement call, colour variation, natural history, and geographic distribution of Proceratophrys caramaschii (Anura, Odontophrynidae). Salamandra, v. 51, p. 103-110, 2015.

24) JUNCÁ, F. A. ; NAPOLI, M. F. ; NUNES, I. ; MERCES, E. A. ; ABREU, R. O. . A New Species of The Scinax ruber Clade (Anura, Hylidae) From The Espinhaço Range, Northeastern Brazil. Herpetologica, v. 71, p. 299-309, 2015.

23) NUNES, I.; SUAREZ, P. ; GORDO, M. ; POMBAL JR., J. P. . A Second Species of Trachycephalus Tschudi (Anura: Hylidae) With A Single Vocal Sac from the Brazilian Amazon. Copeia, v. 2013, p. 634-640, 2013.

22) CAMURUGI, F. ; MERCES, E. A. ; NUNES, I. ; JUNCÁ, F. A. . The tadpole of Scinax strigilatus (Spix, 1824) (Anura: Hylidae). Zootaxa, v. 3686, p. 497-499, 2013.

21) GORDO, M. ; TOLEDO, L. F. ; SUAREZ, P. ; KAWASHITA-RIBEIRO, R. A. ; ÁVILA, R. W. ; MORAIS, D. H. ; NUNES, I. . A new species of milk frog of the genus Trachycephalus Tschudi (anura, hylidae) from the Amazonian Rainforest. Herpetologica, v. 69, p. 466-479, 2013.

20) CARAMASCHI, U. ; NUNES, I. ; NASCIMENTO, L. B. ; CARVALHO JR., R. R. . The type locality of Scinax dolloi (Werner, 1903) (Anura, Hylidae). Zootaxa, v. 3691, p. 299-300, 2013.

19) AMARO, R. C. ; NUNES, I. ; CANEDO, C. ; NAPOLI, M. F. ; JUNCÁ, F. A. ; VERDADE, V. K. ; HADDAD, C. F. B. ; RODRIGUES, M. T. . A molecular phylogeny recovers Strabomantis aramunha Cassimiro, Verdade and Rodrigues, 2008 and Haddadus binotatus (Spix, 1824) (Anura: Terrarana) as sister taxa. Zootaxa, v. 3741, p. 569-582, 2013.

18) POMBAL JR., J. P. ; MENEZES, V. A. ; FONTES, A. F. ; NUNES, I. ; ROCHA, C. F. D. ; VAN SLUYS, M. . A second species of the casque-headed frog genus Corythomantis (Anura: Hylidae) from Northeastern Brazil, the distribution of C. greeningi, and comments on the genus. Boletim do Museu Nacional, Nova Série Zoologia, v. 530, p. 1-14, 2012.

17) NUNES, I.; KWET, A. ; POMBAL JR., J. P. . Taxonomic revision of the Scinax alter species complex (Anura: Hylidae). Copeia, v. 2012, p. 554-569, 2012.

16) JUNCÁ, F. A. ; NAPOLI, M. F. ; CEDRAZ, J. ; NUNES, I. . Acoustic characteristics of the advertisement and territorial calls of Phyllodytes tuberculosus Bokermann, 1966 (Amphibia: Anura: Hylidae). Zootaxa, v. 2012, p. 87-88, 2012.

15) CRUZ, C. A. G. ; NUNES, I. ; JUNCA, F. A. . Redescription of Proceratophrys cristiceps (Müller, 1883) (Amphibia, Anura, Odontophrynidae), with description of two new species without eyelid appendages from Northeastern Brazil. South American Journal of Herpetology, v. 7, p. 110-122, 2012.

14) NUNES, I.; POMBAL JR., J. P. . A New snouted treefrog of the speciose genus Scinax Wagler (Anura, Hylinae), from the Northeastern Brazil. Herpetologica, v. 67, p. 80-88, 2011.

13) CRUZ, C. A. G. ; NUNES, I. ; DE LIMA, M. G. . A new Scinax Wagler belonging to the S. catharinae clade (Anura: Hylidae) from the State of Alagoas, northeastern Brazil. Zootaxa, v. 2011, p. 18-26, 2011.

12) NUNES, I.; COSTA, F. R. . Hide and seek?: diurnal refuge and camouflage of two anurans from the Atlantic Forest of Northeastern Brazil. Herpetology Notes, v. 4, p. 431-433, 2011.

11) NUNES, I.; POMBAL JR., J. P. . A new Scinax Wagler (Amphibia, Anura, Hylidae) from the Atlantic Rain Forest remains of southern State of Bahia, North-eastern Brazil. Amphibia-Reptilia, v. 31, p. 347-353, 2010.

10) NUNES, I.; CANEDO, C. ; CARVALHO JR., R. R. . Advertisement call and geographic distribution of Elachistocleis piauiensis Caramaschi & Jim, 1983 (Amphibia, Microhylidae), with Notes on the presence of post-commissural gland in the genus. South American Journal of Herpetology, v. 5, p. 30-34, 2010.

09) MACIEL, D. B. ; NUNES, I. . A new species of four-eyed frog genus Pleurodema Tschudi, 1838 (Anura: Leiuperidae) from the rock meadows of Espinhaço range, Brazil. Zootaxa, v. 2640, p. 53-61, 2010.

08) NUNES, I.; CARVALHO JR., R. R. ; PEREIRA, E. G. . A new species of Scinax Wagler (Anura: Hylidae) from Cerrado of Brazil. Zootaxa, v. 2514, p. 24-34, 2010.

07) JUNCÁ, F. A. ; NUNES, I. . A new species of marsupial frog of the genus Gastrotheca Fitzinger (Anura: Amphignathodontidae) from the State of Bahia, Northeastern Brazil. Zootaxa, v. 1907, p. 61-68, 2008.

06) NUNES, I.; SANTIAGO, R. S. ; JUNCÁ, F. A. . Advertisement calls of four hylid frogs from the State of Bahia, Northeastern Brasil (Amphibia, Anura, Hylidae). South American Journal of Herpetology, v. 2, p. 89-96, 2007.[89:acofhf];2

05) NUNES, I.. Hypsiboas faber (Blacksmith treefrog). Predation. Herpetological Review, v. 38, p. 186-186, 2007.

04) NUNES, I.; FUSINATTO, L. A. ; CRUZ, C. A. G. . The tadpole and advertisement call of Sphaenorhynchus palustris Bokermann, 1996 (Amphibia, Anura, Hylidae). South American Journal of Herpetology, v. 2, p. 123-128, 2007.[123:ttaaco];2

03) PIMENTA, B. V. S. ; NUNES, I. ; CRUZ, C. A. G. . Notes on the of the poorly known phyllomedusine frog Hylomantis aspera Peters, 1872 (Anura, Hylidae). South American Journal of Herpetology, v. 2, p. 206-214, 2007.[206:notpkp];2

02) ​​NUNES, I.; JUNCÁ, F. A. . Advertisement calls of three Leptodactylid Frogs in the State of Bahia, Northeastern Brazil (Amphibia, Anura, Leptodactylidae), with consideration of their taxonomic status. Arquivos do Museu Nacional, Rio de Janeiro, v. 64, n.2, p. 151-157, 2006.


01) SILVA-FILHO, I. S. N.; JUNCÁ, F. A. . Evidence of full species status of the neotropical leaf-frog Phyllomedusa burmeisteri bahiana (A. Lutz, 1925) (Amphibia, Anura, Hylidae). Zootaxa, v. 1113, p. 51-64, 2006.

Book Chapters

01) SILVA, J. L. ; NUNES, I. ; COSTA, F. R. . Capítulo 1. Mas, afinal, o que é um pterossauro?. In: Fabiana Rodrigues Costa. (Org.). A saga de um paleontólogo: como se estudam os fósseis? 01 ed. Santo André: EdUFABC, 2021, p. 17-25.

02) MACIEL, B. S. ; NUNES, I. . Capítulo 4. Curadoria pós-preparação. In: Fabiana Rodrigues Costa. (Org.). A saga de um paleontólogo: como se estudam os fósseis? 01 ed. Santo André: EdUFABC, 2021, p. 61-69.

03) COSTA, F. R. ; NUNES, I. . Capítulo 6. Ossos e Mais Ossos e o que fazer com eles. In: Fabiana Rodrigues Costa. (Org.). A saga de um paleontólogo: como se estudam os fósseis? 01 ed. Santo André: EdUFABC, 2021, p. 81-93.

​​Other Publications (Scientific Notes and Non Peer-reviewed Material)

COSTA, F. R. ; NUNES, I. Seis meses sem o Museu Nacional. UFABC Divulga Ciência, v. 2, n.3, p. 3, 2019 (Divulgação Científica).

​​NUNES, I. REVIEW: Arteaga, A.; Bustamante, L. & Guayasamin, J. (2013) The Amphibians and Reptiles of Mindo. Life in the cloudforest. Universidad Tecnológica Idoamérica, Quito. 258pp. Herpetologia Brasileira, v.3, n.1, p. 10-11, 2014 (Resenha de Livro).

NUNES, I. ; TINOCO, M. S. ; WARD, R. IUCN Report: Scinax cretatus Nunes & Pombal, 2011. Cambridge: IUCN, 2014 (IUCN Red List of Threatened Species Report).

NUNES, I. REVIEW: Köhler, Gunther. 2011. Amphibians of Central America. Herpeton Verlag, Offenbach. 379 pp. ISBN 3-936180-33-4. Herpetologia Brasileira, v.2, n.1, p. 05-06, 2013 (Resenha de Livro).

COSTA, F. R. ; NUNES, I. REVIEW: Gallo, V., H. M. A. Silva, P. M. Brito & F. J. Figueiredo. Paleontologia de Vertebrados: Relações entre América do Sul e África. Rio de Janeiro: Interciência, 1ª Edição, 2012. 336 p.. Herpetologia Brasileira, v.2, n.1, p. 06-07, 2013 (Resenha de Livro).

​​​​NUNES, I. ; BOWLES, P. IUCN Report: Gastrotheca flamma Juncá & Nunes, 2008. Cambridge: IUCN, 2010 (IUCN Red List of Threatened Species Report).

NUNES, I. ; BOWLES, P. IUCN Report: Scinax tigrinus Nunes, Carvalho & Pereira, 2010. Cambridge: IUCN, 2010 (IUCN Red List of Threatened Species Report).

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